
during the sea-level lowstands. For example, along much of
the east and southern coasts of the United States, Holocene
barrier-lagoon complexes lie only a short distance seaward of
similar deposits of late Pleistocene age (Figure C57d; Demarest
and Kraft, 1987). This leads to enormous stratigraphic com-
plexity, especially in areas of low sediment supply. Sea-level
oscillations with a similar period but much reduced amplitude
probably occurred during “greenhouse” times such as the
Cretaceous. Coastal deposits formed under these conditions
may not show as much complexity because a smaller number
of significant unconformities will be present (Figure C58a).
Summary
The coastal zone is arguably the most complex environment on
earth because of the many independent factors that influence its
characteristics. Pronounced spatial gradients in the intensity of
the main physical processes (river currents, waves, and tidal
currents) create complex assemblages of facies, which are then
stacked in complex stratigraphic successions because of the
migration of the shoreline in response to sea-level change and
sediment addition or removal. The geologic setting and climate
have less direct, but nevertheless important, influences on the
nature of coastal environments. As a result of the interplay of
these many factors, coastal deposits are a rich source of infor-
mation about past environmental conditions.
Robert W. Dalrymple and Kyungsik Choi
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Cross-references
Astronomical Theory of Climate Change
Carbonates, Cool Water
Carbonates, Warm Water
Cretaceous Warm Climates
Deltaic Sediments, Climate Records
Glacial Eustasy
Greenhouse (warm) Climates
Icehouse (cold) Climates
Sea Level Change, Last 250 Million Years
Sea Level Change, Post-Glacial
Sea Level Change, Quaternary
Sea Level Indicators
Sedimentary Indicators of Climate Change
COCCOLITHS
Coccoliths are an important group of microfossils much used in
paleoceanographic studies. They are minute (typically 1–10 µ)
calcite platelets that are produced by unicellular planktonic
algae, coccolithophores. In life, the individual coccolithophore
cell is typically almost entirely covered by coccoliths, forming
an exoskeleton or coccosphere. In the fossil record, coccoliths
are often accompanied by nannoliths, i.e., similar sized calcite
fossils of uncertain affinities (e.g., discoasters, nannoconids).
Geologists study coccoliths and nannoliths together and refer
to them as calcareous nannofossils.
Coccolithophores belong to the algal division (phylum)
Haptophyta, which also includes many non-calcifying taxa
(e.g., Phaeocystis, Prymnesium and Pavlova). Haptophytes
are distinguished from other algae by thei r possession of
two smooth flagella and a unique third flagellum-like organelle,
the haptonema. Molecular genetic research has confirmed that
the Haptophyta are a discrete clade that probably diverged
from other protists in the Early Paleozoic and subsequently
COCCOLITHS 187