on the complete organism. Thus, ducks grow webbed feet so that they can
swim.
Descartes rejected the use of teleological explanation in physics or
biology. Final causation, he maintained, implied in the agent a knowledge
of the end to be pursued; but such knowledge could only exist in minds.
The explanation of every physical movement and activity must be mech-
anistic; that is, it must be given in terms of initial, not final, conditions, and
those conditions must be stated in desc riptive, not evaluative, terms.
Descartes offered no good argument for his contention, and his thesis
ruled out straightforward gravitational attraction no less than the Aristo-
telian cosmic ballet. Moreover, Descartes was wrong to think that teleo-
logical explanation must involve conscious purpose: whatever Aristotle
may have thought about the heavenly bodies, he never believed that an
earthworm, let alone a falling pebble, was in possession of a mind.
It was not Descartes, but Newton and Darwin, who dealt the serious
blows to Aristotelian teleology, by undermining, in different ways, its two
constituent elements. Newtonian gravity, no less than Aristotelian motion,
provides an explanation by reference to a terminus: gravity is a centripetal
force, a force ‘by which bodies are drawn, or impelled, or in any way tend,
towards a point as to a centre’. But Newton’s explanation is fundamentally
different from Aristotle’s in that it involves no suggestion that it is in any
way good for a body to arrive at the centre to which it tends.
Darwinian explanations in terms of natural selection, on the other hand,
resemble Aristotle’s in demanding that the terminus of the process to be
explained, or the complexity of the structure to be accounted for, shall be
something that is beneficial to the relevant organism. But unlike Aristotle,
Darwin explains the processes and the structures, not in terms of a pull by
the final state or perfected structure, but in terms of the pressure of the
initial conditions of the system and its environment. The red teeth and red
claws involved in the struggle for existence were, of course, in pursuit of
a good, namely the survival of the individual organism to which they
belonged; but they were not in pursuit of the ultimate good that is to be
explained by selection, namely, the survival of the fittest species. It is thus
that the emergence of particular sp ecies in the course of evolution could be
explained not only without appeal to a conscious designer, but without
evoking teleology at all.
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