
The Urochordates are animals that literally have a ‘‘cord’’ (chord) within
their ‘‘tail’’ (uro-). By this it is meant that these creatures have a swimming
larval stage with a notochord in its tail section. The main examples of
Urochordates are the sea squirts. Interestingly enough, the adult stage of
the sea squirt has a body shaped like a ‘‘U,’’ and it remains anchored to a
rock at the bottom of the sea. At one end lies an intake siphon, which sucks in
seawater and filters out plankton and other tiny creatures during feeding.
And at the other end lies an output siphon, which squirts or shoots out a jet of
water whenever the animal is bothered by a predator.
The Cephalochordates are literally a group of ‘‘cord-heads’’ (cephalo-)!
The implication of this odd name is displayed in Figure 12.2 (B) for the
lancelets (LANS-lits). The lancelets are slender, fish-like marine animals
that lie partially buried in the sand under shallow water. Their thin, tapered
body is pointed at both ends, making it look much like a small spear or ‘‘little
lance.’’ Because it has no skull, the lancelet doesn’t have a brain, either! Thus,
the notochord extends all the way up into its head, officially making it a
‘‘cord-head’’ (Cephalochordate). The lancelet is a type of suspension feeder,
meaning that it feeds on small particles suspended in seawater, which it draws
into its mouth with the help of waving, hair-like tentacles (TEN-tuh-kuls).
Both sea squirts and lancelets have notochords in their backs during at
least part of their life cycles, but they are still officially invertebrates. Many
biologists think that they are important evolutionary ‘‘bridging species’’
between the other invertebrates (which don’t even have a notochord) and
the true vertebrates (whose primitive notochord has been replaced by a
bony vertebral column).
THE VERTEBRATES – CHORDATES WITH A ‘‘JOINTED
BACKBONE’’
In contrast to the Urochordates and Cephalochordates, the Vertebrata (ver-
teh-BRA-tuh) or vertebrates are chordates with both a jointed (segmented)
backbone and a brain case or cranium (KRAY-nee-um) (see Figure 12.3). You
may recall (Chapter 11) that the ‘‘jointed backbones’’ are technically called
the vertebrae. The cranium is just a formal name for the ‘‘skull’’ (crani)
‘‘present’’ (-um) at the top of the vertebral column. Owing to their high
degree of cephalization (concentration of sensory, motor, and other nerve
functions within the head region), vertebrates have a brain in their cranium.
The vertebrates generally have a notochord only as part of their embryonic
stage of development. As their bodies mature, the notochord is replaced by a
vertebral column (linked series of jointed vertebrae).
[13:25 13/6/03 N:/4058 LAYMAN.751/4058-Alltext.3d] Ref: 4058 Layman: Biology Demystified All-text Page: 194 1-388
2, Web
3, Web
PART 3 Five Kingdoms of Life, plus Viruses
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