
17.2 Cemetery Organization and Brood Care 385
emergence of a more complex behavior of cluster formation. It was further observed
that cemeteries are sited around spatial heterogeneities [77, 563].
A similar behavior is observed in many ant species, such as Leptothorax unifasciatus,
in the way that the colony cares for the brood [77, 912, 913]. Larvae are sorted in such
a way that different brood stages are arranged in concentric rings. Smaller larvae are
located in the center, with larger larvae on the periphery. The concentric clusters are
organized in such a way that small larvae receive little individual space, while large
larvae receive more space.
While these behaviors are still not fully understood, a number of studies have resulted
in mathematical models to simulate the clustering and sorting behaviors. Based on
these simulations, algorithms have been implemented to cluster data, to draw graphs,
and to develop robot swarms with the ability to sort objects. This section discusses
these mathematical models and algorithms. Section 17.2.1 discusses the basic ant
colony clustering (ACC) model, and a generalization of this model is given in Sec-
tion 17.2.2. A minimal ACC approach is summarized in Section 17.2.3.
17.2.1 Basic Ant Colony Clustering Model
The first algorithmic implementations that simulate cemetery formation were inspired
by the studies of Chr´etien of the ants Lasius niger [127]. Based on physical experi-
ments, Chr´etien derived the probability of an ant dropping a corpse next to an n-cluster
as being proportional to 1 − (1 − p)
n
for n ≤ 30, where p is a fitting parameter [78].
Ants cannot, however, precisely determine the size of clusters. Instead, the size of
clusters is determined by the effort to transport the corpse (the corpse may catch on
other items, making the walk more difficult). It is likely that the corpse is deposited
when the effort becomes too great.
On the basis of of Chr´etien’s observations, Deneubourg et al. [200] developed a model
to describe the simple behavior of ants. The main idea is that items in less dense areas
should be picked up and dropped at a different location where more of the same type
exist. The resulting model is referred to as the basic model.
Assuming only one type of item, all items are randomly distributed on a two-
dimensional grid, or lattice. Each grid-point contains only one item. Ants are placed
randomly on the lattice, and move in random directions one cell at a time. After each
move, an unladen ant decides to pick up an item (if the corresponding cell has an
item) based on the probability
P
p
=
γ
1
γ
1
+ λ
2
(17.43)
where λ is the fraction of items the ant perceives in its neighborhood, and γ
1
> 0.
When there are only a few items in the ant’s neighborhood, that is λ<<γ
1
,thenP
p
approaches 1; hence, objects have a high probability of being picked up. On the other
hand, if the ant observes many objects, that is λ>>γ
1
, P
p
approaches 0, and the
probability that the ant will pick up an object is small.