
Environmental Encyclopedia 3
Savanna
Saprophyte
A saprophyte is an organism that survives by consuming
nutrients from dead and decaying plant and animal material,
that is, organic matter. Saprophytes include
fungi
, molds,
most bacteria, actinomycetes, and a few plants and animals.
Saprophytes contain no chlorophyll and are, therefore, un-
able to produce food through
photosynthesis
, the conver-
sion of chemical compounds into energy when light is pres-
ent. Organisms that do not produce their own food,
heterotrophs, obtain nutrients from surrounding sources,
living or dead. For saprophytes, the source is non-living
organic matter. Saprophytes are known as
decomposers
.
They absorb nutrients from forest floor material, reducing
complex compounds in organic material into components
useful to themselves, plants, and other
microorganisms
.
For example, lignin, one of three major materials found in
plant cell walls, is not digestible by plant-eating animals
or useable by plants unless broken down into its various
components, mainly complex sugars. Certain saprophytic
fungi are able to reduce lignin into useful compounds. Sapro-
phytes thrive in moist, temperate to tropical environments.
Most require oxygen to live. North American plant sapro-
phytes include truffles, Indian Pipe (Monotropa uniflora),
pinedrops (Pterospora andromedea), and snow orchid (Cepha-
lanthera austinae), all of which feed on forest floor litter in
the northeast forests.
[Monica Anderson]
SARA
see
Superfund Amendments and
Reauthorization Act (1986)
Savanna
A savanna is a dry grassland with scattered trees. Most
ecologists agree that a characteristic savanna has an open or
sparse canopy with 10–25% tree cover, a dominant ground
cover of annual and perennial grasses, and less than 20 in
(50 cm) of rainfall per year. Greatly varied environments,
from open deciduous forests and parklands, to dry, thorny
scrub, to nearly pure
grasslands
, can be considered savan-
nas. At their margins these communities merge, more or
less gradually, with drier prairies or with denser, taller forests.
Savannas occur at both tropical and temperate latitudes and
on all continents except
Antarctica
. Most often savannas
occupy relatively level, or sometimes rolling, terrain. Charac-
teristic savanna soils are dry, well-developed ultisols, oxisols,
and alfisols, usually basic and sometimes lateritic. These soils
develop under savannas’ strongly seasonal rainfall regimes,
1251
with extended dry periods that can last up to 10 months.
Under natural conditions an abundance of insect, bird, rep-
tile, and mammal
species
populate the land.
Savanna shrubs and trees have leathery, sometimes
thorny, and often small leaves that resist
drought
, heat,
and intense sunshine normally found in this
environment
.
Savanna grasses are likewise thin and tough, frequently
growing in clumps, with seasonal stalks rising from longer-
lived underground roots. Many of these plants have oily or
resinous leaves that burn intensely and quickly in a fire.
Extensive root systems allow most savanna plants to exploit
moisture and nutrients in a large volume of
soil
. Most sa-
vanna trees stand less than 32 ft (10 m) tall; some have a
wide spreading canopy while others have a narrower, more
vertical shape. Characteristic trees of African and South
American savannas include acacias and miombo
(Brachystegia spp.). Australian savannas share the African
baobab, but are dominated by eucalyptus species. Oaks char-
acterize many European savannas, while in North America
oaks, pines, and aspens are common savanna trees.
Because of their extensive and often nutritious grass
cover, savannas support extensive populations of large herbi-
vores. Giraffes,
zebras
, impalas, kudus, and other charis-
matic residents of African savannas are especially well-
known. Savanna herbivores in other regions include North
American
bison
and elk and Australian kangaroos and wal-
labies. Carnivores—lions, cheetahs, and jackals in Africa,
tigers
in Asia,
wolves
and pumas in the Americas—histori-
cally preyed upon these huge herds of grazers. Large running
birds, such as the African ostrich and the Australian emu,
inhabit savanna environments, as do a plethora of smaller
animal species. In the past century or two many of the
world’s native savanna species, especially the large carnivores,
have disappeared with the expansion of human settlement.
Today ranchers and their livestock take the place of many
native grazers and carnivores.
Savannas owe their existence to a great variety of con-
vergent environmental conditions, including temperature
and precipitation regimes, soil conditions, fire frequency,
and
fauna
. Grazing and browsing activity can influence the
balance of trees to grasses. Fires, common and useful for
some savannas but rare and harmful in others, are an influen-
tial factor in these dry environments. Precipitation must be
sufficient to allow some tree growth, but where rainfall is
high some other factors, such as grazing, fire, or soil
drain-
age
needs to limit tree growth. Human activity also influ-
ences the occurrence of these lightly-treed grasslands. In
some regions recent expansion of ranches, villages, or agricul-
ture have visibly extended savanna conditions. Elsewhere
centuries or millennia of human occupation make natural
and
anthropogenic
conditions difficult to distinguish. Be-
cause savannas are well suited to human needs, people have